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In the basal state PNPLA2 has weak hydrolytic activity which is greatly increased by the co-lipase alpha/beta hydrolase domain-containing protein 5 (ABHD5, also known as CGI-58) by greater than 20-fold (30)

Importantly, a mutation in PNPLA3 (rs738409), I148M, is the single greatest genetic risk factor for the development of FLD (64-67), however its precise function in neutral lipid metabolism is controversial (68, 69)

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