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A senior Japan Railway official said the group has never forgotten the fact that smokers have been helping pay off the JNR debt

Weevils have elongated snouts that differentiate them from other insects

E-cigarettes and cigars are left out of the bill because Dr Creagan believes they are significantly safer for smokers than regular cigarettes - although the National Cancer Institute warns that "all tobacco products are harmful and cause cancer"

Cigarette butts are composed of thousands of cellulose acetate fibres and, although biodegradable, take years to disappear from the environment

The researchers hypothesize that this area modulates value